Professional and Biographical Information


Ph.D. (Biochemistry and Molecular Biology), Harvard University (1975)
M.A. (Chemistry), Yale University (1968)
B.S. (Chemistry), Yale University (1968)
A.M. ( honorary), Amherst College (1994)

Teaching Interests

For many years I taught Molecular Genetics and Biochemisty, two courses mirroring my research interests.  More recently I have taught the introductory course Molecules, Genes, and Cells, an advanced seminar on regulatory RNA molecules (DNA makes RNA), and the first year seminar Genes, Genomes, and Society.

Scholarly and Professional Activities

Associate Faculty Member, Program in Molecular & Cellular Biology, University of Massachusetts, Amherst, Massachusetts, 1984-present

Visiting Scientist, Imperial Cancer Research Fund, London, 1990-1991

Visiting Scientist, NIDDKD, NIH, 1995

Acting Dean of the Faculty, 1999-2000

Visiting Scientist, Columbia University Medical School, 2000-2001

Program Director, Amherst College HHMI Science Education Award, 2004-2009

See also: Research Interests and Selected Publications